# STOCKHOLM 1.0 #=GF ID Lar_N #=GF AC PF09861.12 #=GF DE Lactate racemase N-terminal domain #=GF PI DUF2088; #=GF AU COGs; #=GF AU Finn RD;0000-0001-8626-2148 #=GF AU Sammut SJ;0000-0003-4472-904X #=GF AU El-Gebali S;0000-0003-1378-5495 #=GF SE COGs (COG3875) #=GF GA 23.30 23.30; #=GF TC 23.30 23.30; #=GF NC 23.20 23.20; #=GF BM hmmbuild HMM.ann SEED.ann #=GF SM hmmsearch -Z 61295632 -E 1000 --cpu 4 HMM pfamseq #=GF TP Domain #=GF CL CL0471 #=GF RN [1] #=GF RM 24710389 #=GF RT Lactate racemase is a nickel-dependent enzyme activated by a #=GF RT widespread maturation system. #=GF RA Desguin B, Goffin P, Viaene E, Kleerebezem M, Martin-Diaconescu #=GF RA V, Maroney MJ, Declercq JP, Soumillion P, Hols P; #=GF RL Nat Commun. 2014;5:3615. #=GF DR INTERPRO; IPR018657; #=GF DR SO; 0000417; polypeptide_domain; #=GF CC This is the N-terminal domain found in Lactate racemase enzymes #=GF CC EC:5.1.2.1. LarA is a nickel-dependent lactate racemase from the #=GF CC larA-E operon of Lactobacillus plantarum, whereas LarB, LarC and #=GF CC LarE are accessory proteins required for the incorporation of Ni #=GF CC in the lactate racemase apoprotein. LarA contains 18 #=GF CC beta-strands and 16 alpha-helices arranged in a novel fold #=GF CC composed of two domains of similar size. This entry represents #=GF CC the N-terminal domain of LarA and related proteins [1]. #=GF SQ 11615 X1L7B1.1/11-152 ----.---.-.---..-..-..-..-....-.....-..-.....-..-...-...--..........--..-...-....-....-..-..P....D.E..I...K...A...SI...S...N.P..I................G...I............P......P...............I....R....E......L....A......K......G.....K.................K....E............V...........V.I......IF.D..D.....M....T..R.....V.....TR.V...........................A.K.I.M.P.F.V.L.E.E..L..A.A..A..G..I..P....D...N..R......I..R..F..I.V...A..L.....G.C...H...G..A..L.D......R......L......D......F......V......K.......K..........L......G..........E.......E......V......V...A..R.F...............................P.V...Y.N..H..N.......-.........-...........P.........F..........A.......N......C........T........Y.............V.............G.................T..............T.............S.....................T................yK..............T..............K........V...Y..V..NE.E..V..M..........G..........C......D..L..KIAIGSVVPH...G....G...A...G...F.....E...G.....-.....-.....-...------...----..--------..---------- X1L7B1.1/151-287 ----.---.-.---..-..-..-..-....-.....-..-.....-..-...-...--..........--..-...-....-....-..-..-....-.-..-...-...-...--...-...-.-..-................-...-............-......-...............-....-....-......-....-......E......G.....K.................K....E............V...........V.I......IF.D..D.....M....T..R.....V.....TR.V...........................A.K.I.M.P.F.V.L.E.E..L..A.A..A..G..I..P....D...N..R......I..R..F..I.V...A..L.....G.L...H...S..T..M.W......R......Q......H......F......V......K.......K..........L......G..........E.......E......V......V...A..R.F...............................P.V...Y.N..H..N.......-.........-...........P.........F..........Y.......N......C........T........Y.............V.............G.................T..............T.............S.....................T................yK..............T..............R........V...Y..A..NE.E..V..M..........K..........C......D..L..KIAIGSVVPH...P....M...S...G...F.....G...G.....G.....G.....K...IIMPGV...ASFE..TID-----..---------- A0A6I6ET65.1/8-204 YGNG.AID.V.SIA..E..E..D..L....L.....G..V.....I..D...S...SV..........EV..D...N....K....S..E..E....E.V..I...R...K...AL...E...N.P..I................G...S............N......K...............L....K....E......L....V......H......P.....G.................E....K............I...........C.I......VV.S..D.....I....T..R.....A....wQK.M...........................S.V.F.L.P.Y.I.I.E.E..L..N.G..A..G..I..K....D...E..D......I..I..F..I.S...A..T.....G.S...H...R..G..Q.T......E......E......E......H......K......S.......I..........L......G..........E.......E......L......Y...K..R.F...............................T.I...I.D..H..E.......C.........T...........D.........K..........E.......N......L........T........Y.............I.............G.................T..............T.............S.....................F.................G..............T..............P........I...S..I..NK.I..A..A..........E..........C......D..H..IVLTGAIVFH...L....L...V...G...W.....G...G.....G.....K.....K...SLLPGI...SSYE..TIMANHAL..TL-------- A0A1I4GMY0.1/8-204 YGKD.KLH.F.TIP..E..E..N..L....L.....G..V.....L..T...S...DA..........PQ..N...E....Q....T..E..A....E.I..I...K...T...AL...A...N.P..I................G...S............R......R...............L....A....E......I....V......K......P.....G.................E....Q............I...........C.I......VI.S..D.....I....T..R.....S....wQR.M...........................G.D.Y.L.P.Y.L.V.E.E..L..Q.Q..A..G..V..K....A...E..D......I..M..F..L.S...A..T.....G.S...H...R..K..Q.T......Q......A......E......H......N......T.......L..........L......G..........E.......K......L......A...G..K.F...............................K.V...I.D..H..V.......A.........K...........D.........E..........A.......S......L........V........Y.............L.............G.................E..............T.............S.....................Y.................G..............T..............P........V...K..I..NK.L..A..A..........E..........A......D..H..IILTGAIVFH...L....L...A...G...W.....G...G.....G.....K.....K...SVLPGI...AGYE..SIMANHAM..SL-------- A0A1M6MNB5.1/8-206 YGKT.KID.V.SIP..Q..E..N..L....L.....G..I.....I..A...S...NK..........EI..R...N....K....T..E..E....E.V..V...L...E...AL...D...K.P..I................E...S............N......K...............L....R....E......I....V......N......K.....G.................D....K............V...........C.I......VI.S..D.....I....T..R.....A....wQK.M...........................N.F.Y.L.P.Y.I.V.E.E..L..E.K..A..D..I..R....E...E..D......I..I..F..L.C...A..T.....G.T...H...R..K..Q.T......K......K......E......H......G......I.......L..........L......G..........E.......K......L......A...K..R.F...............................Q.V...I.D..H..D.......C.........M...........D.........K..........D.......N......L........V........Y.............L.............G.................T..............T.............S.....................F.................G..............T..............P........V...S..I..NK.I..A..L..........E..........C......D..H..IIITGAIVYH...D....M...A...G...W.....G...G.....G.....R.....K...SILPGI...SSYE..SIMANHAL..ALN-K----- D9S157.1/10-208 YGRG.MMT.L.KIP..A..K..N..V....L.....G..V.....L..K...A...RD..........VP..A...V....P....D..E..K....E.A..V...V...E...AV...R...N.P..I................G...S............P......P...............L....S....G......L....V......K......K.....G.................D....S............V...........V.I......IA.S..D.....I....T..R.....S.....VR.N...........................S.R.V.I.P.V.L.L.E.E..L..R.E..S..G..I..P....Y...E..D......I..T..V..V.V...A..T.....G.T...H...R..G..H.T......E......Q......E......L......E......Y.......M..........F......G..........R.......E......V......L...G..R.V...............................R.V...V.D..H..N.......C.........R...........D.........R..........E.......S......L........V........Y.............L.............G.................T..............T.............S.....................S.................G..............T..............P........V...W..V..NR.L..V..A..........G..........A......D..K..VIITGGIVYH...S....M...A...G...F.....G...G.....G.....R.....K...AVCPGV...SGYE..TIQHNHKL..MLNPP----- A0A5S5AYL5.1/10-206 YGRG.TMT.L.KIP..A..R..N..V....L.....G..V.....L..E...T...QA..........VP..A...V....P....D..E..K....E.A..V...V...E...AV...R...N.P..T................G...S............P......P...............L....S....E......L....V......K......K.....G.................D....S............V...........V.V......IA.S..D.....I....T..R.....N.....VR.N...........................P.V.V.I.P.T.I.L.E.E..L..R.E..S..G..I..P....Y...E..D......I..T..V..V.V...A..T.....G.T...H...R..G..H.T......R......Q......E......L......E......F.......M..........F......G..........R.......E......V......L...E..R.V...............................R.V...V.D..H..N.......C.........R...........D.........R..........E.......S......L........V........Y.............L.............G.................V..............T.............S.....................R.................G..............T..............P........V...W..V..NR.L..V..A..........E..........A......D..K..VIITGGIVYH...S....M...A...G...F.....G...G.....G.....R.....K...AVCPGV...SGYE..TIQYNHKL..ML-N------ A0A2W4JYF9.1/10-209 YGRK.VVK.L.QIP..E..D..N..L....L.....G..V.....L..K...T...GN..........IP..A...V....D....D..E..R....Q.A..V...V...N...AL...R...N.P..I................A...S............P......P...............L....H....E......I....V......K......A.....G.................D....K............V...........V.I......IA.S..D.....I....T..R.....N.....AR.N...........................P.V.V.I.P.A.V.I.D.E..L..K.G..A..G..V..K....P...E..D......M..T..I..V.V...A..T.....G.T...H...R..G..H.S......R......E......E......L......E......Y.......L..........F......G..........A.......D......V......I...E..Q.V...............................R.V...V.D..H..D.......S.........R...........N.........R..........D.......N......L........V........H.............I.............G.................T..............T.............S.....................R.................G..............T..............P........V...W..I..NR.L..V..A..........E..........A......D..K..VIITGGIVYH...S....M...A...G...F.....G...G.....G.....R.....K...AICPGV...SSFE..TIQHNHKL..MLNPPE---- A0A4R2KLP6.1/8-210 YGRG.TVD.V.SIP..K..K..N..L....L.....K..V.....I..E...S...NS.........sKE..K...S....K....T..Q..E....E.V..I...I...D...AL...A...N.P..I................A...S............A......R...............L....K....Q......L....V......H......Q.....G.................E....K............V...........C.V......VI.S..D.....V....T..R.....L....wQK.M...........................S.M.Y.L.P.Y.I.V.E.E..L..N.N..G..G..V..K....D...E..D......I..L..F..I.S...A..T.....G.S...H...R..E..Q.T......K......E......E......H......T......L.......L..........L......G..........E.......K......L......S...E..R.F...............................E.V...I.N..H..N.......C.........H...........D.........E..........E.......N......L........T........Y.............L.............G.................K..............T.............R.....................F.................G..............T..............P........V...M..V..NK.K..A..L..........E..........C......D..H..IILTGAIVFH...L....L...A...G...W.....G...G.....G.....K.....K...SVLPGI...CSYE..SIMKNHAL..SL-SPNAG-- A0A1T5J2H1.1/8-208 YGKE.KVN.L.CLN..E..E..N..L....M.....G..V.....I..E...S...NK..........ID..N...K....K....S..E..E....E.V..I...L...N...AL...E...N.P..I................D...S............P......K...............L....C....D......I....V......N......E.....G.................E....K............I...........C.I......VI.S..D.....I....T..R.....A....wQK.M...........................G.A.Y.L.H.Y.I.V.D.E..L..N.K..G..G..V..S....D...E..D......I..V..F..L.S...A..T.....G.S...H...R..K..Q.S......K......E......E......H......D......I.......L..........L......G..........E.......K......L......A...D..R.F...............................E.V...I.D..H..D.......C.........L...........D.........K..........E.......N......L........V........Y.............L.............G.................K..............T.............S.....................F.................G..............T..............P........V...E..I..NR.I..A..M..........G..........C......D..H..IILTGAIVFH...L....L...A...G...W.....G...G.....G.....K.....K...SILPGI...SSYD..SIMKNHSM..SL-NPEM--- A0A6M0T0S3.1/8-213 YHKK.MIE.I.EIE..N..K..N..L....V.....G..I.....L..E...S...KAe........dYK..R...N....L....S..Q..E....E.I..V...E...A...AL...D...N.P..I................Q...C............E......K...............L....E....D......L....V......K......G.....K.................K....N............V...........V.I......IT.S..D.....H....T..R.....P.....VP.S...........................K.I.T.M.P.I.L.L.R.R..I..R.E..-..V..N..P....D...I..D......I..K..I..L.I...A..T.....G.F...H...R..E..S.T......K......E......E......L......I......N.......K..........F......G..........E.......E......I......V...K..N.E...............................N.I...V.M..H..V.......S.........T...........D.........Y..........S.......T......L........V........K.............V.............G.................K..............L.............P.....................S.................G..............G..............E........L...I..L..NK.L..A..M..........E..........A......E..L..LIAEGFIESH...F....F...A...G...F.....S...G.....G.....R.....K...SILPGI...ASAK..TIMANHCG.eFIASPNARTG A0A6M0XV96.1/8-213 YHKK.MIE.I.EIE..N..K..N..L....V.....G..I.....L..E...S...KAe........dYK..R...N....L....S..Q..E....E.I..V...E...A...AL...D...N.P..I................Q...C............E......K...............L....E....D......L....V......K......G.....K.................K....N............V...........V.I......IT.S..D.....H....T..R.....P.....VP.S...........................K.I.T.M.P.I.L.L.R.R..I..R.E..-..V..N..P....D...I..D......I..K..I..L.I...A..T.....G.F...H...R..E..S.T......K......E......E......L......I......N.......K..........F......G..........E.......E......I......V...K..N.E...............................N.I...V.M..H..V.......S.........T...........D.........Y..........S.......T......L........V........K.............V.............G.................K..............L.............P.....................S.................G..............G..............E........L...I..L..NK.L..A..M..........E..........A......E..L..LIAEGFIESH...F....F...A...G...F.....S...G.....G.....R.....K...SILPGI...ASAK..TIMANHCG.eFIASPNARTG A0A402BP13.1/9-206 YGHE.EML.L.KID..E..E..H..I....L.....Q..V.....I..E...S...NQ..........VT..F...E....K....S..E..S....E.M..I...R...A...AL...A...N.P..I................G...S............P......R...............L....S....E......L....V......K......S.....G.................E....R............V...........C.I......VI.P..D.....I....T..R.....A....wQR.T...........................D.L.Y.L.P.L.L.V.E.E..L..A.Q..G..G..V..Q....D...E..D......I..L..F..I.C...A..L.....G.T...H...R..P..Q.T......A......E......E......H......R......L.......L..........L......G..........P.......E......L......A...D..R.F...............................T.V...I.D..H..D.......C.........H...........D.........R..........T.......N......L........V........Y.............C.............G.................E..............T.............S.....................F.................G..............T..............P........V...W..L..NK.Q..A..M..........E..........C......D..H..LVLTGGIVYH...F....L...A...G...W.....S...G.....G.....K.....K...YVLPGI...SSYE..TVMANHAL..SL-N------ A0A1S8LIW6.1/7-204 YGDL.KLE.I.DVD..E..K..N..V....I.....G..V.....I..K...S...NK..........FG..T...T....K....S..E..E....E.V..I...K...D...AI...E...N.P..I................D...S............A......R...............L....K....E......L....V......H......E.....G.................E....T............V...........C.I......VV.P..D.....V....T..R.....A....wQR.T...........................D.K.Y.L.P.K.L.V.E.E..I..N.A..G..G..V..K....D...E..D......I..I..F..L.S...A..T.....G.T...H...R..S..Q.T......E......E......E......H......E......K.......I..........L......G..........K.......E......L......S...K..R.F...............................K.I...I.D..H..D.......C.........L...........D.........K..........N.......N......L........V........Y.............L.............G.................K..............T.............T.....................Y.................G..............T..............P........V...S..I..NK.I..A..L..........D..........C......D..H..IVITGGIVYH...F....L...V...G...F.....S...G.....G.....K.....K...TILPGI...ASYE..SVMANHAM..SLS------- M1MT13.1/8-205 YGQG.EMK.A.LIE..E..K..N..L....L.....E..V.....I..E...S...NV..........LR..Q...D....R....T..E..A....E.I..I...S...E...AL...L...N.P..I................D...S............K......R...............L....K....D......I....V......H......E.....D.................E....R............V...........C.I......VI.S..D.....I....T..R.....S....wQK.P...........................H.E.F.L.Y.K.I.I.E.E..L..N.L..G..G..V..K....D...E..N......I..I..F..L.C...A..Q.....G.T...H...R..K..Q.T......K......E......E......H......E......I.......L..........L......G..........E.......E......L......S...K..R.F...............................E.V...Y.D..H..D.......C.........F...........D.........E..........E.......N......L........V........Y.............L.............G.................D..............T.............S.....................Y.................G..............T..............P........V...K..V..NK.I..A..V..........E..........A......D..H..VVITGAIVYH...F....L...V...G...W.....S...G.....G.....K.....K...SILPGI...SSFK..TISANHSL..SLN------- J7SV28.1/8-204 YGKD.SIK.F.SLS..E..E..N..L....L.....G..V.....I..D...R...NE..........WN..L...D....K....T..E..E....E.I..I...K...E...SI...E...N.P..I................C...S............A......P...............L....K....E......L....V......H......E.....G.................E....K............I...........C.I......VI.P..D.....I....T..R.....A....wQR.S...........................S.V.Y.L.P.Y.I.V.E.E..I..K.K..G..G..V..K....D...E..D......I..I..F..L.S...S..T.....G.T...H...R..E..Q.T......E......E......E......H......K......I.......L..........I......G..........E.......K......L......Y...N..N.Y...............................K.V...V.D..H..I.......S.........T...........K.........K..........E.......D......L........V........Y.............V.............G.................K..............T.............S.....................Y.................N..............T..............P........V...M..I..NK.L..A..L..........E..........C......D..H..VILTGAIIYH...F....L...V...G...Y.....S...G.....G.....K.....K...AILPGI...SSFE..TVMANHSL..SL-------- A0A6M0T4L6.1/8-204 YGKD.SIK.F.SLP..E..E..N..I....L.....G..V.....I..D...R...NK..........WN..L...D....K....T..E..E....E.I..I...K...D...SI...E...N.P..I................C...S............A......P...............L....K....E......L....V......H......E.....G.................E....K............I...........C.I......VI.P..D.....I....T..R.....A....wQR.S...........................S.V.Y.L.P.Y.I.V.E.E..I..K.K..G..G..V..K....D...E..D......I..I..F..L.S...S..T.....G.T...H...R..K..Q.T......E......E......E......H......K......I.......L..........I......G..........E.......K......L......Y...K..N.Y...............................K.I...I.D..H..V.......S.........T...........K.........K..........E.......D......L........V........Y.............V.............G.................K..............T.............S.....................Y.................N..............T..............P........I...M..I..NK.L..A..L..........E..........C......D..H..VILTGAIIYH...F....L...V...G...Y.....S...G.....G.....K.....K...AILPGI...SSFE..TVMANHAL..SL-------- A0A6M0Y1Q5.1/8-204 YGKD.SIK.F.SLP..E..E..N..I....L.....G..V.....I..D...R...NK..........WN..L...D....K....T..E..E....E.I..I...K...D...SI...E...N.P..I................C...S............A......P...............L....K....E......L....V......H......E.....G.................E....K............I...........C.I......VI.P..D.....I....T..R.....A....wQR.S...........................S.V.Y.L.P.Y.I.V.E.E..I..K.K..G..G..V..K....D...E..D......I..I..F..L.S...S..T.....G.T...H...R..K..Q.T......E......E......E......H......K......I.......L..........I......G..........E.......K......L......Y...K..N.Y...............................K.I...I.D..H..V.......S.........T...........K.........K..........E.......D......L........V........Y.............V.............G.................K..............T.............S.....................Y.................N..............T..............P........I...M..I..NK.L..A..L..........E..........C......D..H..VILTGAIIYH...F....L...V...G...Y.....S...G.....G.....K.....K...AILPGI...SSFE..TVMANHAL..SL-------- A0A0M1IVR8.1/8-204 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A0A6B3Y327.1/8-209 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M5E396.1/8-213 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A0A662QZE8.1/2-199 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A0A2N8Z6N0.1/32-237 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A0A151ANH5.1/8-213 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