# STOCKHOLM 1.0 #=GF ID DDAH_eukar #=GF AC PF19420.2 #=GF DE N,N dimethylarginine dimethylhydrolase, eukaryotic #=GF AU Chuguransky S;0000-0002-0520-0736 #=GF GA 23.0 23.0 #=GF NC 22.9 22.9 #=GF TC 23.0 23.0 #=GF SE Pfam-B_5784 (release 5.2) and Pfam-B_1850 (release 5.5) #=GF BM hmmbuild HMM.ann SEED.ann #=GF SM hmmsearch -Z 61295632 -E 1000 --cpu 4 HMM pfamseq #=GF TP Family #=GF CL CL0197 #=GF RN [1] #=GF RM 11504612 #=GF RT A novel superfamily of enzymes that catalyze the modification of #=GF RT guanidino groups. #=GF RA Shirai H, Blundell TL, Mizuguchi K; #=GF RL Trends Biochem Sci 2001;26:465-468. #=GF RN [2] #=GF RM 14675764 #=GF RT Prediction of the structure and function of AstA and AstB, the #=GF RT first two enzymes of the arginine succinyltransferase pathway of #=GF RT arginine catabolism. #=GF RA Shirai H, Mizuguchi K; #=GF RL FEBS Lett 2003;555:505-510. #=GF DR SCOP; 1jdw; fa; #=GF DR SO; 0100021; polypeptide_conserved_region; #=GF CC This family contains N(G),N(G)-dimethylarginine #=GF CC dimethylaminohydrolases (DDAH) from eukaryotes. It also includes #=GF CC arginine deiminases and DDAH from prokaryotes. These enzymes are #=GF CC involved in arginine metabolism and belong to the #=GF CC amidinotransferase (AT) superfamily as they share the alpha/beta #=GF CC propeller fold, which includes structurally important residues #=GF CC (buried hydrophobic residues, buried hydrophilic residues #=GF CC hydrogen-bonded with mainchain groups and the hallmark of three #=GF CC consecutive buried Gly residues near the C-terminus, conserved #=GF CC among its members [1,2]. #=GF SQ 5495 Q8RGP8.1/21-309 FTFN.EETAVNNHYQKK............DN.K...P...I....Q...E........I........Q.NN.AL...T.E..FDNM....V.EKL..K...K...T...G...I..D..V...K....V...M.......QD......T.K...K.......P.......H.......T....P......D.......S......IFPNN....W.FS.T................HY.....S.....N.....T.V.....V....L.Y.P..M.F.AE.N....R..RL.E..R...T..D....N.LY.DYFDET........--.D.NL.....N..V.VDYS......S..L.E..N.......E..N.I.F.LEG.....T.....GS......L.......V......L........D...R..K........N...K....K........A.........Y........C.........S..........L..........S.........E..........R.ANE....K....L....L...D...I.F...C......K...D.A.....G.......Y.....K.....KI.A.F.H..S.Y..Q..........T.......V....D.E.K......R.K.P.....I..Y..HTN........V...M.M.A.M.........G.E...N...Y......A........I......L.........C........A.D....S.I........D..N..L..K.E.....R..ENV..I.RE.....LK.....N.....D.D.....KEIVYI..S..E..Y..Q..V.....E...H..F..LG...N.TIEL-.......INN.EN.VNIC..vMSATA..YS.V.L.TD..E.Q...K.N.I..I.EK......Y..D..V..I..V..PVDV.HT.IERYGGGSARCMIAELF A0A0P0M2M7.1/8-298 FGFN.EETAANNYFQQR............TV.A...A...G....E...T........V........Q.AC.AL...K.E..FRGM....V.DKL..R...A...N...G...I..E..V...I....A...V.......QD......T.P...I.......P.......H.......T....P......D.......S......IFPNN....W.VS.F................HD.....D.....G.....T.V.....V....L.Y.P..M.F.AE.N....R..RA.E..R...R..A....D.IL.QRIESE........GF.S.IR.....R..K.VDYT......G..F.E..E.......E..N.R.F.LEG.....T.....GG......I.......I......L........D...R..V........N...R....Y........A.........Y........A.........A..........L..........S.........Q..........R.VDE....A....L....F...G...R.F...C......K...E.Q.....N.......F.....S.....PV.S.F.H..A.Y..Q..........T.......V....N.G.Q......R.L.P.....I..Y..HTN........V...M.M.C.V.........A.T...D...Y......A........V......V.........C........L.D....S.I........D..D..V..E.E.....R..EQV..C.HV.....LR.....Q.....T.G.....KEIIAI..S..E..E..Q..M.....H...H..F..AG...N.MLQVE.......NGK.GA.LFLV...MSQSA..YD.S.L.DE..G.Q...L.Q.R..L.SK......Y..N..A..L..L..PMPI.PT.IETLGGGSARCMMAEIF A6L2I9.1/17-307 FGFN.EETAANNYFQQR............TV.A...A...G....E...T........V........Q.AC.AL...K.E..FRGM....V.DKL..R...A...N...G...I..E..V...I....A...V.......QD......T.P...I.......P.......H.......T....P......D.......S......IFPNN....W.VS.F................HD.....D.....G.....T.V.....V....L.Y.P..M.F.AE.N....R..RA.E..R...R..A....D.IL.QRIESE........GF.S.IR.....R..K.VDYT......G..F.E..E.......E..N.R.F.LEG.....T.....GG......I.......I......L........D...R..V........N...R....Y........A.........Y........A.........A..........L..........S.........Q..........R.VDE....A....L....F...G...R.F...C......K...E.Q.....N.......F.....I.....PV.S.F.H..A.Y..Q..........T.......V....N.G.Q......R.L.P.....I..Y..HTN........V...M.M.C.V.........A.T...D...Y......A........V......V.........C........L.D....S.I........D..D..V..E.E.....R..EQV..C.HV.....LR.....Q.....T.G.....KEIIAI..S..E..E..Q..M.....H...H..F..AG...N.MLQVE.......NGK.GA.LFLV...MSQSA..YD.S.L.DE..G.Q...L.Q.R..L.SK......Y..N..A..L..L..PMPI.PT.IETLGGGSARCMMAEIF R9HQE7.1/7-297 FGFN.EETAANNYFQQR............TV.A...A...G....E...T........V........Q.AC.AL...K.E..FRGM....V.DKL..R...A...N...G...I..E..V...I....A...V.......QD......T.P...I.......P.......H.......T....P......D.......S......IFPNN....W.VS.F................HD.....D.....G.....T.V.....V....L.Y.P..M.F.AE.N....R..RA.E..R...R..A....D.IL.QRIESE........GF.S.IR.....R..K.VDYT......G..F.E..E.......E..N.R.F.LEG.....T.....GG......I.......I......L........D...R..V........N...R....Y........A.........Y........A.........A..........L..........S.........Q..........R.VDK....A....L....L...G...R.F...C......K...E.Q.....N.......F.....S.....PV.S.F.H..A.Y..Q..........T.......V....N.G.Q......R.L.P.....I..Y..HTN........V...M.M.C.V.........A.T...D...Y......A........V......V.........C........L.D....S.I........D..D..V..E.E.....R..EQV..C.HV.....LR.....Q.....T.G.....KEIIAI..S..E..E..Q..M.....H...H..F..AG...N.MLQVE.......NGK.GA.LFLV...MSQSA..YD.S.L.DE..G.Q...L.Q.R..L.SK......Y..N..A..L..L..PMPI.PT.IETLGGGSARCMMAEIF A0A3A5T2L6.1/7-297 FGFN.EETAANNYFQQR............TV.A...A...G....E...T........V........Q.AC.AL...K.E..FRGM....V.DKL..R...A...N...G...I..E..V...I....A...V.......QD......T.Q...I.......P.......H.......T....P......D.......S......IFPNN....W.VS.F................HD.....D.....G.....T.V.....V....L.Y.P..M.F.AE.N....R..RA.E..R...R..A....D.IL.QRIESE........GF.C.IR.....R..K.VDYT......G..F.E..E.......E..N.R.F.LEG.....T.....GG......I.......I......L........D...R..V........N...R....Y........A.........Y........A.........A..........L..........S.........Q..........R.VDE....A....L....F...G...R.F...C......K...E.Q.....N.......F.....I.....PV.S.F.H..A.Y..Q..........T.......V....N.G.Q......R.L.P.....I..Y..HTN........V...M.M.C.V.........A.T...D...Y......A........V......V.........C........L.D....S.I........D..D..V..E.E.....R..EQV..C.HV.....LR.....Q.....T.G.....KEIIAI..S..E..E..Q..M.....H...H..F..AG...N.MLQVE.......NGK.GA.LFLV...MSQSA..YD.S.L.DE..G.Q...L.Q.R..L.SK......Y..N..A..L..L..PMPI.PT.IETLGGGSARCMMAEIF X7RVS6.1/17-305 FAFN.EQTAVNNHYQKR............DN.K...P...V....Q...E........I........Q.NK.AL...I.E..FDNM....V.EKL..K...K...V...G...I..D..V...K....V...M.......QD......T.K...E.......P.......H.......T....P......D.......S......IFPNN....W.FS.T................HY.....S.....N.....T.V.....V....L.Y.P..M.F.AE.N....R..RL.E..R...T..D....T.IY.DYFD--........RV.D.NL.....N..I.VDYS......S..L.E..K.......E..N.I.F.LEG.....T.....GS......L.......V......L........D...R..K........N...K....K........A.........Y........C.........S..........L..........S.........E..........R.ADE....K....L....L...D...I.F...C......E...D.A.....G.......Y.....K.....KI.A.F.H..S.Y..Q..........T.......V....D.G.K......R.K.P.....I..Y..HTN........V...M.M.A.M.........G.E...N...Y......A........I......L.........C........A.D....S.I........D..D..L..K.E.....R..EKV..I.RE.....LE.....S.....D.G.....KEIVYI..S..E..Y..Q..V.....E...H..F..LG...N.AIEL-.......INN.EN.VKIC..vMSATA..YS.V.L.TD..E.Q...K.N.I..I.EK......Y..D..V..I..V..PVDV.HT.IEKYGGGSARCMIAELF A0A069SLJ2.1/7-297 FGFN.EETAANNYFQQR............TV.A...A...G....E...T........V........Q.AC.AL...K.E..FRGM....V.DKL..R...A...N...G...I..E..V...I....A...V.......QD......T.P...I.......P.......H.......T....P......D.......S......IFPNN....W.VS.F................HD.....D.....G.....T.V.....V....L.Y.P..M.F.AE.N....R..RA.E..R...R..A....D.IL.QRIESE........GF.C.IR.....R..K.VDYT......G..F.E..E.......E..N.R.F.LEG.....T.....GG......I.......I......L........D...R..V........N...R....Y........A.........Y........A.........A..........L..........S.........Q..........R.VDE....A....L....F...G...R.F...C......K...E.Q.....N.......F.....S.....PV.S.F.H..A.Y..Q..........T.......V....N.G.Q......R.L.P.....I..Y..HTN........V...M.M.C.V.........A.T...D...Y......A........V......V.........C........L.D....S.I........D..D..V..E.E.....R..EQV..C.HV.....LR.....Q.....T.G.....KEIIAI..S..E..E..Q..M.....H...H..F..AG...N.MLQVE.......NGK.GA.LFLV...MSQSA..YD.S.L.DE..G.Q...L.Q.R..L.SK......Y..N..A..L..L..PMPI.PT.IETLGGGSARCMMAEIF A3HWB9.2/7-297 FGFN.TETAANNFYQQM............DD.R...S...G....K...E........I........Q.EL.AR...K.E..FDGF....V.ALL..R...D...Q...R...V..N..V...E....V...V.......ED......T.E...N.......P.......V.......K....T......D.......A......VFPNN....W.FS.T................HS.....D.....G.....K.L.....I....M.Y.P..M.F.SP.N....R..RL.E..R...R..K....D.LI.EMLMHK........GY.Q.VS.....D..I.LDLS......F..F.E..N.......D..G.Q.Y.LEG.....T.....GS......M.......V......M........D...H..E........S...K....T........I.........F........A.........C..........Y..........S.........E..........R.THP....F....P....L...Q...Y.V...A......E...L.L.....D.......Y.....K.....VV.G.F.H..S.V..Q..........E.......I....E.G.V......Q.S.P.....I..Y..HTN........V...M.M.H.I.........G.R...D...L......A........V......V.........C........L.D....S.I........P..K..A..S.E.....K..QLV..Q.RT.....LT.....N.....N.G.....KKVIPI..T..P..K..Q..K.....F...N..F..AG...N.MLELS.......NDG.GE.KFTI...MSQTA..YD.S.L.NP..G.Q...I.Q.L..I.EK......Y..T..T..I..I..SPQI.PT.IEKLGGGSARCMIAEIF A0A133NEZ3.1/21-309 FAFN.EQTAVNNHYQKL............DN.K...P...A....Q...E........I........Q.NN.AL...I.E..FDNM....V.EKL..K...K...I...G...I..D..V...R....V...M.......QD......T.K...E.......P.......H.......T....P......D.......S......IFPNN....W.FS.T................HY.....S.....N.....T.V.....V....L.Y.P..M.F.AE.N....R..RL.E..R...T..D....N.LY.DYFDKA........--.D.DL.....N..I.VDYS......N..L.E..K.......E..N.I.F.LEG.....T.....GA......L.......V......L........D...R..K........N...K....K........A.........Y........C.........S..........L..........S.........E..........R.ANE....K....L....L...D...I.F...C......K...D.A.....G.......Y.....K.....KI.A.F.H..S.Y..Q..........T.......V....D.E.K......R.K.P.....I..Y..HTN........V...M.M.A.M.........G.E...N...Y......V........I......L.........C........A.D....S.I........D..N..L..E.E.....R..ENV..I.RE.....LK.....N.....D.G.....KEIVYI..T..E..Y..Q..V.....E...H..F..LG...N.TIEL-.......VNN.EN.VNIC..vMSATA..YS.V.L.TN..E.Q...K.N.I..I.EK......Y..D..V..I..V..PVDV.HT.IERYGGGSARCMIAELF F7L1Y1.1/17-305 FAFN.EQTAVNNHYQKL............DN.K...P...A....Q...E........I........Q.NN.AL...I.E..FDNM....V.EKL..K...K...I...G...I..D..V...R....V...M.......QD......T.K...E.......P.......H.......T....P......D.......S......IFPNN....W.FS.T................HY.....S.....N.....T.V.....V....L.Y.P..M.F.AE.N....R..RL.E..R...T..D....S.LY.DYFDKA........--.D.DL.....N..I.VDYS......N..L.E..K.......E..N.I.F.LEG.....T.....GA......L.......V......L........D...R..K........N...K....K........A.........Y........C.........S..........L..........S.........E..........R.ANE....K....L....L...D...I.F...C......E...D.A.....G.......Y.....K.....KI.A.F.H..S.Y..Q..........T.......V....A.G.K......R.K.P.....I..Y..HTN........V...M.M.A.M.........G.E...N...Y......A........I......L.........C........A.D....S.I........D..N..L..E.E.....R..ENV..I.RE.....LE.....N.....D.G.....KEIVYI..T..E..N..Q..V.....E...H..F..LG...N.TIEL-.......VNN.EN.VNIC..vMSATA..YS.V.L.TN..E.Q...K.N.I..I.EK......Y..D..V..I..V..PVDV.HT.IERYGGGSARCMIAELF C3RFG3.2/7-297 FGFN.EETAANNYFQQR............TV.A...A...G....E...T........V........Q.AC.AL...K.E..FRGM....V.DKL..R...A...N...S...I..E..V...I....V...V.......QD......T.P...I.......P.......H.......T....P......D.......S......IFPNN....W.VS.F................HD.....D.....G.....T.V.....V....L.Y.P..M.F.AE.N....R..RA.E..R...R..A....D.IL.QRIESE........GF.S.IR.....R..K.VDYT......G..F.E..E.......E..N.R.F.LEG.....T.....GG......I.......I......L........D...R..V........N...R....Y........A.........Y........A.........A..........L..........S.........Q..........R.VDE....A....L....L...G...R.F...C......K...E.Q.....N.......F.....I.....PV.P.F.R..A.Y..Q..........T.......V....N.G.Q......R.L.P.....I..Y..HTN........V...M.M.C.V.........A.T...D...Y......A........V......V.........C........L.D....S.I........D..D..V..E.E.....R..EQV..C.HI.....LR.....Q.....T.G.....KEIIAI..S..E..E..Q..M.....H...H..F..AG...N.MLQVE.......NGK.GA.LFLV...MSQSA..YD.S.L.DE..G.Q...L.Q.R..L.SK......Y..N..A..L..L..PMPI.PT.IETLGGGSARCMMAEIF I9R7F4.1/7-297 FGFN.EETAANNYFQQR............TV.A...A...G....E...T........V........Q.AC.AL...K.E..FRGM....V.DKL..R...A...N...G...I..E..V...I....V...V.......QD......T.P...I.......P.......H.......T....P......D.......S......IFPNN....W.VS.F................HD.....D.....G.....T.V.....V....L.Y.P..M.F.AE.N....R..RA.E..R...R..A....D.IL.QRIESE........GF.S.IR.....R..K.VDYT......G..F.E..E.......E..N.R.F.LEG.....T.....GG......I.......I......L........D...R..V........N...R....Y........A.........Y........A.........A..........L..........S.........Q..........R.VDE....A....L....L...G...R.F...C......K...E.Q.....N.......F.....I.....PV.P.F.R..A.Y..Q..........T.......V....N.G.Q......R.L.P.....I..Y..HTN........V...M.M.C.V.........A.T...D...Y......A........V......V.........C........L.D....S.I........D..D..V..E.G.....R..EQV..C.HI.....LR.....Q.....T.G.....KEIIAI..S..E..E..Q..M.....H...H..F..AG...N.MLQVE.......NGK.GA.LFLV...MSQSA..YD.S.L.HE..D.Q...L.Q.R..L.SK......Y..N..A..L..L..SMPI.PT.IETLGGGSARCMMAEIF H1HHS8.1/17-305 FAFN.EQTAVNNHYQKL............DN.K...P...A....Q...E........I........Q.NN.AL...I.E..FDNM....V.EKL..K...K...I...G...I..D..V...R....V...M.......QD......T.K...E.......P.......H.......T....P......D.......S......IFPNN....W.FS.T................HY.....S.....N.....T.V.....V....L.Y.P..M.F.AE.N....R..RL.E..R...T..D....N.LY.DYFDKA........--.D.DL.....N..I.VDYS......N..L.E..K.......E..N.I.F.LEG.....T.....GA......L.......V......L........D...R..K........N...K....K........A.........Y........C.........S..........L..........S.........E..........R.ANE....K....L....L...D...I.F...C......E...D.A.....G.......Y.....K.....KI.A.F.H..S.Y..Q..........T.......V....D.G.K......R.K.P.....I..Y..HTN........V...M.M.A.M.........G.E...N...Y......A........I......L.........C........T.D....S.I........D..N..L..E.E.....R..ENV..I.RE.....LK.....N.....D.G.....KEIVYI..S..E..F..Q..V.....D...H..F..LG...N.TIEL-.......VNN.EN.VNIC..vMSATA..YS.V.L.TN..E.Q...K.N.I..I.EK......Y..D..V..I..V..PVDV.HT.IEKYGGGSARCMIAELF D6BHY5.1/17-305 FAFN.EQTAVNNHYQKL............DN.K...P...A....Q...E........I........Q.NN.AL...I.E..FDNM....V.EKL..K...K...I...G...I..D..V...R....V...M.......QD......T.K...E.......P.......H.......T....P......D.......S......IFPNN....W.FS.T................HY.....S.....N.....T.V.....V....L.Y.P..M.F.AE.N....R..KL.E..R...T..D....N.LY.DYFDKA........--.D.DL.....N..I.VDYS......N..L.E..K.......E..N.I.F.LEG.....T.....GA......L.......V......L........D...R..K........N...K....K........A.........Y........C.........S..........L..........S.........E..........R.ANE....K....L....L...D...I.F...C......E...D.A.....G.......Y.....K.....KI.A.F.H..S.Y..Q..........T.......V....G.G.K......R.K.P.....I..Y..HTN........V...M.M.A.M.........G.E...N...Y......A........I......L.........C........A.D....S.I........D..N..L..K.E.....R..ENV..I.KE.....LE.....N.....D.G.....KEIVYI..S..E..Y..Q..V.....E...H..F..LG...N.AIEL-.......INN.EN.VNIC..vMSATA..YS.V.L.TN..E.Q...K.N.I..I.EK......Y..D..V..I..V..SVDV.HT.IERYGGGSARCMIAELF C1DBM8.1/17-309 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